Ceratolobus Blume ex Schult. & Schult.f.
  • Syst. Veg. 7: lxxx (1830) 


Notes: Distribution: Pen. Thailand to W. Malesia

General Description

Slender to moderate, clustered, spiny, climbing, pleonanthic, dioecious, rattan palms. Stem eventually becoming bare, with long internodes and conspicuous nodal scars, white mucilage sometimes exuding from cut surfaces. Leaves of mature climbing stems cirrate, pinnate; leaf sheath tubular, armed with spines and/or spicules, frequently organised into whorls, and often with abundant indumentum; knee present, sometimes rather weakly developed; ocrea inconspicuous; flagellum absent; petiole present or absent, if present, flat adaxially, rounded abaxially, armed with spines and sometimes with spicules; cirrus and distal part of rachis armed with regular groups of grapnel spines on the abaxial surface; leaflets relatively few, linear to lanceolate and entire, or rhomboid and praemorse, concolourous or discolourous, regularly arranged or grouped; emerging leaf pink-tinged. Inflorescences axillary but adnate to the internode and sheath of the following leaf, sessile and erect or pendulous on a long slender, unarmed or spiny peduncle, the whole inflorescence much shorter than the leaves; staminate inflorescence branching to 3 orders, pistillate to 2 orders, prophyll persistent, membranous to subwoody, flattened-tubular with lateral wings and a terminal beak, entirely enclosing the inflorescence, opening at anthesis by 2 narrow, lateral slits in the beak, this remaining the only access to the flowers during anthesis and young fruiting stage, prophyll unarmed or rarely armed with scattered spines, frequently bearing caducous indumentum, prophyll often splitting longitudinally in fruit, very rarely falling completely, staminate and pistillate inflorescences indistinguishable without splitting the prophyll; peduncular bracts absent; rachis bracts inconspicuous, tubular, with triangular limbs, each (and the prophyll) subtending a first-order branch, usually adnate to the axis for a short distance above the bract node. Staminate flowers borne singly, subdistichously, rather distant from each other, each subtended by a membranous triangular bract and a 2-keeled, prophyllar bracteole, the latter ± forming a cushion beneath the flower; calyx tubular, with 3 short triangular lobes; petals 3, boat-shaped, valvate, briefly joined basally; stamens 6, borne at the base of the corolla, filaments short, fleshy, anthers linear, latrorse; pistillode trifid, minute. Pollen ellipsoidal, bi-symmetric; apertures equatorially disulcate; ectexine tectate or semi-tectate, finely or coarsely perforate, or foveolate-reticulate, aperture margins similar or finer; infratectum columellate; longest axis 22–30 µm [3/6]. Pistillate flowers borne with a sterile staminate flower and 2 similar 2-keeled prophyllar bracteoles, in a cup formed by the subtending bract. Sterile staminate flower like the fertile but usually rather distorted by close packing and with empty anthers and borne on a short to long stalk. Pistillate flowers larger than the staminate; calyx tubular, with 3 short, triangular lobes; corolla partially divided into 3, valvate, triangular lobes; staminodes 6, epipetalous, flattened; gynoecium incompletely trilocular, triovulate, globose, or ellipsoidal, covered in scales, stigmas 3, fleshy, recurved, borne on a short style, ovule basally attached, anatropous. Fruit l-seeded, globose to ellipsoidal, stigmatic remains apical, epicarp covered in vertical rows of reflexed scales, mesocarp becoming thin and papery as fruit ripens, endocarp not differentiated. Seed attached basally, globose to ellipsoidal, with thin or thick, sour or sweet sarcotesta and homogeneous or ruminate endosperm; embryo basal. Germination adjacent-ligular; eophyll with 4–6 praemorse or entire, crowded leaflets displayed in a fan. Cytology: 2n = 26.

Diagnostic Description

Slender clustering climbing palms, found in South Thailand, Malay Peninsula, Borneo, Sumatra and Java, immediately distinguished by the single large bract (the prophyll) that covers the entire inflorescence and opens by two apical slits.

Morphology

Leaf (Tomlinson 1961), root (Seubert 1996a).

Biology

All species are found in lowland and hill tropical rain forest and do not occur above 1000 m altitude; all are usually confined to dipterocarp forest but Ceratolobus subangulatus also occurs in heath forest in Sarawak, where it is the most conspicuous rattan in some facies. The pollination ecology of the extraordinary closed inflorescence deserves further study.

Distribution

Six species confined to the perhumid areas of the Sunda Shelf (Malay Peninsula [2], Sumatra [4], Borneo [3], Java [2]).

Uses

Species of Ceratolobus appear to have weak, not durable canes and for this reason are scarcely ever utilised.

Common Names

Rattan, rotan.

Distribution Map

 
  • Native distribution
Found in
  • Asia-Tropical Indo-China Thailand
  • Malesia Borneo
  • Jawa
  • Malaya
  • Sumatera

  Bibliography

  • 1 J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008
  • 2 Govaerts, R. & Dransfield, J. (2005). World Checklist of Palms: 1-223. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • 3 Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. Continental Publishing, Deurne.

 Information From

Palmweb - Palms of the World Online
http://www.palmweb.org
Palmweb 2011. Palmweb: Palms of the World Online. Published on the internet http://www.palmweb.org. Accessed on 21/04/2013
  • A Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
  • B http://creativecommons.org/publicdomain/zero/1.0
eMonocot
http://e-monocot.org
eMonocot. (2010, 1st November). Retrieved Wednesday, 8th February, 2012, from http://e-monocot.org.
  • C Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
World Checklist of Selected Plant Families
http://apps.kew.org/wcsp/
WCSP 2014. 'World Checklist of Selected Plant Families. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://apps.kew.org/wcsp/ Retrieved 2011 onwards
  • D See http://kew.org/about-kew/website-information/legal-notices/index.htm You may use data on these Terms and Conditions and on further condition that: The data is not used for commercial purposes; You may copy and retain data solely for scholarly, educational or research purposes; You may not publish our data, except for small extracts provided for illustrative purposes and duly acknowledged; You acknowledge the source of the data by the words "With the permission of the Trustees of the Royal Botanic Gardens, Kew" in a position which is reasonably prominent in view of your use of the data; Any other use of data or any other content from this website may only be made with our prior written agreement.